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How did Megatron begin?

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The name is a title , kind of like king or emperor . The Fallen was once Megatronus Prime , I would guess this as the origin of the name . As for the origin of the person , I am sure that many decepticon leaders have carried this title , think of it like the name ' darth ' from the star wars series , you must become the apprentice of a certain someone to earn this title , in the case of the transformers-verse , The Fallen took an apprentice and bestowed the name upon him . Best guess .

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